Heat flow data from the Polish basin show a sharp change in the transition from the East European Craton (EEC) and Teisseyre–Tornquist Zone (TTZ) in the north-east to the accreted terranes in the south west (Paleozoic Platform). The analysis of this data and numerical modelling of the crustal temperatures show evidence of extensive crustal–mantle warming in the area between the Sudetes to the south and the Trans-European Suture Zone to the north. The change in heat flow is 100% when compared with values for the EEC. Heat flow in the anomalous zone is also higher than in the Sudetes. The axis of the anomaly is aligned with the Dolsk Fault and Variscan deformation front. Low crustal/mantle temperatures derived from the relationship between temperature and Pn velocities (more than 8.2 and as high as 8.4 km/s) are at odds with high crustal temperatures calculated from surface heat flow, seismic velocity based heat generation models and thermal conductivity. High heat flow (Variscan platform) and related high temperatures of the crust coincide with small crustal thickness (30–35 km). The opposite is the case for the low heat flow EEC (45–50 km). High heat flow above thin crust and low heat flow above thick crust with no major variation in elevation is supported by a simple isostatic balance model. Crustal heat generation explains part of the high heat flow within the zone with thick meta-sediments reaching down to 20 km depth, however, it is far from explaining high heat flow in Variscan crust and in the transition zone into a cold EEC. 2D numerical models of heat flow based on new seismic data require a contrast of 15 mW/m2 in mantle heat flow. High mantle heat flow (35–40 mW/m2) is likely to occur in the high heat flow zone while cold crust and cold and high-density mantle (mantle heat flow of 20–30 mW/m2) is typical of the EEC. Thermal lithosphere thickness for the craton is 200 km while it is only 100 km in the accreted terranes to the southwest of the TTZ. The TTZ in Poland appears as a relatively cold area.